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Brigita Laime,
Faculty of Biology, University of Latvia
Characteristic
of the seacoasts
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Marine and Seacoast Biodiversity]
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Beaches commonly are
formed by easily washed out sediments (sand, gravel and pebble) and
are directly submerged to regular sea and wind influence. Walking
along the water line from Nida to Ainaži, we can observe sandy,
stony, pebble and gravel beaches. Their width varies from 5-10 to
50-100 meters and even more. Most widespread are sandy beaches where
in comparison with the other beaches higher species and habitat
diversity is observed. Most characteristic plant species are Cakile
baltica, Salsola kali and Honckenya peploides. These
plants commonly grow scattered or in small groups. Most of plant
species grow on low wet beaches where annual plants dominate, like Chenopodium
sp. and Atriplex sp. Well developed beach communities are
in August and September when the species composition has stabilised;
spotted or in some places continuous plant cover is observed.
Differently develop habitats in stony beaches where perennial vascular
plants dominate (Laime 2000).
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Beach
Photo: B.Laime
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Dunes are wind flown sandy ridges. Near the
sea both open dunes and those covered by trees and shrubs occur. In
open dunes two zones are distinguished - primary dunes and grey dunes.
Primary dunes are located most close to the sea and commonly
border with the beach and where sand flooding takes place. Primary
dunes are characteristic only in those sections of coast where sand
accumulation takes place. Total length of primary dunes in Latvia is
about 240 km (Anon. 1993). Development of primary dunes starts from
embryonic dunes that are small up to 50-cm high sandy formations with
scarce vegetation. Characteristic species are Leymus arenarius,
xCalammophila baltica, Honckenya peploides and Elytrigia
x littorea. With the continuation of sand accumulation and plant
cover development these primary dunes develop into foredunes.
Their medium height is 1-3 m
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Dunes
Photo: B.Laime
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although 5-6 m or even higher occur. Mainly there
are coastal sections where is only one foredune, in medium 50 m
wide.In places where sand accumulation processes are more active 2 or
3 foredunes develop that can be 100 and more meter wide. Grasses
dominate on foredunes, the most common are Ammophila arenaria,
Calamagrostis epigeios, Leymus arenarius and Festuca arenaria.
In Latvia foredunes with shrubs are characteristic, mainly with
willows, like Salix viminalis and Salix daphnoides. They
are planted to stabilise dunes or have arrived on their own.
On beaches and primary dunes 282 plant species grow (Laime 2000). Data
obtained from the studies of the botanists of the University of
Latvia, Faculty of Biology from 1989 to 2002. Many species are weeds,
garden escapers and invasive plants. Comparing the flora of beaches
and primary dunes in different sections of the coast it was found out
that more rich is the coast of the Gulf of Rīga, especially between Rīga
and Kolka. Near the Gulf of Rīga 240 species occur, on the coast near
the Baltic Sea from Nida to Kolka 113 species. It can be due to the
diversity of coastal habitats and it concerns mainly Kurzeme part of
the Gulf of Rīga.
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A grey dune is a
secondary type of dunes that develops from primary dunes. Grey dunes
are relatively stable. Bryophytes, lichens and low vascular plants as
well as separate trees and shrubs form the plant cover. On grey dunes
in Latvia 163 species of Pteridophyta and Spermatophyta
grow, 30 bryophyte species and 44 lichen species from which 23 are
Latvian Red Data Book species (Laime, Rove 2001). Special attention
must be paid to those species whose habitats are mainly connected with
grey dunes and which have a small distribution area, like Dianthus
arenarius subsp. arenarius.
Grey dunes in Latvia are a rare and specially protected habitat (Anon.
2001) and are distributed unevenly, mainly near the coast of the
Baltic Sea. Width of grey dunes varies from several metres up to 30-50
metres. Near the coast of the Gulf of Rīga the medium width of grey
dunes is 5 metres. Near the coast of the Baltic Sea it is between 5 to
10 metres. Grey dunes are found in Užava (width up to 50 metres) and
in the north of Pāvilosta (width about 100 metres).
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Grey dunes
Photo: B.Laime
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Dune slacks is a part of the coastal
habitat complex and are characteristic for the coastal sections with
low beaches as well in the ecotone zone between foredunes and grey
dunes or dune forest. Rare are dune slacks with calcareous rich fen
species (Laime, Pakalne 2000). Bluffs
are characteristic for the seacoast in Latvia that have developed
under the influence of abrasion and wash away (Ulsts 1998). Especially
interesting are bluffs with springs and calcareous plant species as
well as sandstone outcrops.
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Bluffs
Photo: B.Laime
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For the shallow water part of the Gulf of Rīga typical are wetlands
with tall vascular plant growth where Phragmites australis, Scirpus
tabernaemontani and Bolboschoenus maritimus dominate. Most
of these wetlands border with moist beaches and coastal meadows.
Widest wetlands are located between Ainaži and Salacgrīva as well as
between Bērzciems and Roja.
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Wetlands
Photo: B.Laime
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Flora
Typical plant species on beaches and primary dunes
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Beach
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Cakile baltica
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Atriplex littoralis
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Agrostis stolonifera
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Salsola kali
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Equisetum arvense
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Tussilago farfara
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Juncus articulatus
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Juncus bufonius
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Sagina nodosa
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Primary
dunes
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Ammophila arenaria
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Leymus arenarius
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X Calammophila baltica
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Calamagrostis epigeios
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Festuca arenaria
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Elytrigia x littorea
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Hieracium umbellatum
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Artemisia campestris
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Honckenya peploides
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Salix daphnoides
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Characteristic
species of grey dunes
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Pteridophyta and Spermatophyta
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Arctostaphylos
uva-ursi
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Artemisia
campestris
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Carex
arenaria
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Corynephorus
canescens
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Dianthus
arenarius
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Festuca
sabulosa
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Galium
mollugo
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Galium
verum
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Hieracium
umbellatum
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Jasione
montana
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Koeleria
glauca
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Pulsatilla
pratensis
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Salix
rosmarinifolia
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Sedum
acre
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Thymus
serpyllum
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Epipactis
atrorubens
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Empetrum
nigrum
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Honckenya
peploides
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Viola
canina
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Viola
tricolor
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Bryophytes
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Brachythecium
albicans
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Ceratodon
purpureus
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Dicranum
scoparium
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Hypnum
cupressiforme
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Pleurozium
schreberi
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Racomitrium
canescens
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Racomitrium
ericoides
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Tortella
inclinata
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Tortula
ruralis
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Lichens
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Cetraria
aculeata
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Cetraria
ericetorum
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Cladonia
chlorophea
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Cladonia
fimbriata
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Cladonia
furcata
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Cladonia
glauca
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Cladonia
gracilis
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Cladonia
mitis
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Cladonia
portentosa
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Cladonia
rangiferina
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Cladonia
rangiformis
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Cladonia
verticillata
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Hypogymnia
physodes
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Peltigera
canina
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Peltigera
didactila
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Peltigera
rufescens
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Fauna
Seacoast habitats are important habitats for feeding and migration of
many animal species. In the coastal zone at least half of the Red Data
Book bird species appear, like Tadorna tadorna, Haemotopus
ostralegus, Pluvialis apricaria, Calidris alpina schinzii etc.
Most of them are registered during migration (Ādamsons 1986,
Lipsbergs 1985, Pēterhofs 1984). It concerns also bats. During autumn
migration, for example in Pape ornithological station all 15-bat
species found in Latvia were observed (Pētersons 1997). Sandy beaches
and foredunes are considered as a typical habitat for Bufo calamita.
Up to know several isolated populations are observed at the seacoast
(Bērziņs 1984, 1987, 1988). At the seacoast, especially in places
washed by seawater quite high insect diversity is observed. Several
rare species appear there, like Laphria gibbosa, Laphria flava and
Cicindela maritima. [Top of
page]
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to Marine and Seacoast Biodiversity]
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